Collected Evidence: Collected Evidence: Cease mowing on grassland to allow early succession Sixteen studies evaluated the effects on butterflies and moths of ceasing mowing on grassland to allow early succession. Three studies were in Germany, two were in each of the USA, Spain and the Czech Republic, and one was in each of Switzerland, Poland, Slovakia, Hungary, Japan, Russia and Italy. COMMUNITY RESPONSE (8 STUDIES) Community composition (3 studies): Two replicated, site comparison studies in the Czech Republic and Germany found that the community composition of butterflies and moths in grasslands which had been abandoned for >5 years or an unspecified length of time was similar to grasslands managed by mowing or grazing (results not distinguished). One replicated, controlled, before-and-after study in Spain found that after mowing and grazing was abandoned, over 6 years the butterfly community became dominated by generalist species, and species with fewer generations/year. Richness/diversity (7 studies): Five of seven replicated, site comparison studies in Germany, Poland, Japan, the Czech Republic, Russia and Italy found that grasslands which had been abandoned for 3–13 years, 10–20 years or an unspecified length of time, had a similar species richness of butterflies and burnet moths or all moths to grasslands managed by annual or unspecified frequency mowing, or mown within the last three years (in two studies mowing and grazing were not distinguished). One of these studies also found that grasslands abandoned for more than 50 years had lower species richness than grasslands mowed annually to up to 20 years ago, and another found that grasslands abandoned for 6–13 years had a lower species richness of butterflies than grasslands managed by traditional rotational mowing and burning. One of the studies found that meadows not cut all summer had a higher species richness of butterflies than meadows cut 1–3 times/summer. The other study found that grasslands abandoned for at least 5–20 years had a lower species richness of butterflies than grasslands managed by mowing or grazing (results not distinguished). POPULATION RESPONSE (14 STUDIES) Abundance (14 studies): Four replicated studies (including one randomized, paired, controlled study and three site comparison studies) in Germany, Spain, Slovakia and Hungary found that grasslands which had been abandoned for >1–20 years had a lower abundance of all butterflies or some species of butterfly and caterpillars, than grasslands managed by mowing once or twice per year (in two studies mowing and grazing were not distinguished). Four replicated, site comparison studies (including one paired study) in Germany, the Czech Republic, Russia and Italy found that grasslands which had been abandoned for >3 years, were temporarily abandoned, or were uncut all summer, had a higher abundance of all butterflies, 11 species of butterfly, Scotch argus adults and meadow neb moth caterpillars, than grasslands managed by mowing annually, 1–3 times/summer, or within the last three years. Two replicated studies (including one controlled, before-and-after study and one site comparison study) in Spain and Germany found that grasslands which had been abandoned for 1–6 years or an unspecified time period had a higher abundance of woodland and hedgerow butterflies and burnet moths, but a lower abundance of grassland or farmland species, than grasslands managed by mowing and/or grazing (results not distinguished). One of these studies also found that silver-studded blue went extinct in some abandoned meadows. Three replicated, site comparison studies in the USA and Poland found that in grasslands which had been abandoned for many years or 10 to over 50 years before abundance was mixed depending on butterfly species compared to grasslands managed by grazing or mowing. One replicated, site comparison study in Switzerland found that grasslands which had been abandoned for around six years had a similar abundance of heath fritillary adults and caterpillars to grasslands managed by annual mowing, but that grasslands abandoned for >25 years had a lower abundance of adults and no caterpillars. BEHAVIOUR (2 STUDIES) Use (2 studies): One replicated, paired, site comparison study in Germany found that grassland which had been abandoned for >5 years had a similar occurrence of hoary bell moth caterpillars to grassland managed by mowing. One replicated, site comparison study in Spain found that grizzled skipper and painted lady occurred less frequently, but small pearl-bordered fritillary occurred more frequently, in meadows which had been abandoned for at least 1–2 years than in meadows managed by mowing or grazing (results not distinguished). Collected Evidencehttps%3A%2F%2Fconservationevidencejournal.com%2Factions%2F3957https%3A%2F%2Fconservationevidencejournal.com%2Factions%2F3957Sun, 14 Aug 2022 10:36:48 +0100Collected Evidence: Collected Evidence: Reduce management intensity on permanent grasslands (several interventions at once) Twelve studies evaluated the effects on butterflies and moths of reducing management intensity on permanent grasslands. Seven studies were in Switzerland, three were in the UK, and one was in each of Greece and Germany. COMMUNITY RESPONSE (12 STUDIES) Community composition (2 studies): Two replicated studies (including one controlled study and one site comparison study) in Switzerland found that the composition of butterfly communities differed between low-input and intensively managed grasslands. One of these studies found that low-input grasslands tended to have more butterfly species whose caterpillars feed on a single host plant, have one generation/year and poor dispersal ability. Richness/diversity (11 studies): Six of 10 studies (including five controlled studies and five site comparison studies) in Switzerland, the UK, Greece and Germany found that less intensively managed grasslands had a higher species richness of butterflies and moths than conventionally managed grasslands, although two of these studies only found a difference in one of two years or regions. The other four studies found that less intensively managed grasslands had a similar species richness of butterflies and moths to conventionally managed grasslands. However, one of these studies also found that less intensively managed grassland had more specialist species of moths, and species of conservation concern, than conventionally managed grassland. One before-and-after study in the UK found that after grazing was reduced and chemical application stopped, the species richness of large moths increased. POPULATION RESPONSE (5 STUDIES) Abundance (5 studies): Three of four replicated studies (including two controlled studies and two site comparison studies) in Switzerland, the UK and Germany found that low-input or unfertilized, ungrazed grassland managed with a single cut had a higher abundance of butterflies, micro-moths and declining macro-moths than intensively managed grassland. Two of these studies also found that the abundance of caterpillars and of all macro-moths was similar between less intensively and more intensively managed grasslands. The other study found that less intensively managed grassland had a similar abundance of moths to more intensively managed grassland. One before-and-after study in the UK found that after grazing was reduced and chemical application stopped, the total abundance of large moths and the abundance of five out of 23 butterfly species increased, but the abundance of two butterfly species decreased. BEHAVIOUR (1 STUDY) Use (1 study): One replicated, site comparison study in Germany reported that 24 out of 58 moth species preferred less intensively managed grasslands, but 12 species preferred more intensively managed grasslands. Collected Evidencehttps%3A%2F%2Fconservationevidencejournal.com%2Factions%2F3958https%3A%2F%2Fconservationevidencejournal.com%2Factions%2F3958Sun, 14 Aug 2022 10:37:05 +0100Collected Evidence: Collected Evidence: Reduce grazing intensity on grassland by reducing stocking density Fourteen studies evaluated the effects on butterflies and moths of reducing grazing intensity on grassland by reducing stocking density. Four studies were in the UK, two were in each of Sweden and Germany, one was in each of the USA, Belgium and the Netherlands, Europe and Switzerland. COMMUNITY RESPONSE (8 STUDIES) Richness/diversity (8 studies): Five of eight replicated studies (including two randomized, controlled studies and six site comparison studies) in the UK, Sweden, Germany, the USA and Switzerland found that grasslands grazed with lower stocking densities of sheep, cattle or a mix of sheep, cattle and horses had a greater species richness of adult butterflies, all moths and burnet moths than grassland grazed at higher stocking densities. However, one of these studies also found that butterfly and burnet moth caterpillar species richness was similar at sites with low and high stocking densities. Two of the other studies found that grasslands grazed with lower stocking densities of cattle and horses or unspecified grazing animals had a similar species richness of butterflies and burnet moths to grassland grazed at higher stocking densities. The other study found that, in one of two study years, grasslands grazed with cattle at a low density had lower species richness than grasslands grazed at moderate density. POPULATION RESPONSE (12 STUDIES) Abundance (11 studies): Eight replicated studies (including four controlled studies and four site comparison studies) in the UK, Germany, the USA and Sweden found that grasslands grazed with lower stocking densities of sheep, cattle or both (in one case combined with a later start to grazing) had a greater abundance of all butterflies, butterflies with grass host plants, all moths, burnet moths, their caterpillars or specific species (in two cases as part of combined invertebrate counts) than grasslands grazed at higher stocking densities. The three studies on caterpillars only found a higher abundance at two out of three sites or in earlier or later sampling periods, and one of the studies found that sites with low and high intensity grazing had a similar abundance of butterfly and burnet moth caterpillars. Two replicated, site comparison studies in Sweden and Switzerland found that grasslands grazed with lower stocking densities of cattle and horses or unspecified grazing animals had a similar abundance of butterflies and burnet moths to grassland grazed at higher stocking densities. One review of studies in Europe reported that reducing grazing intensity benefitted 41 out of 67 butterfly species of conservation concern, but did not distinguish between reducing stocking density and seasonal removal of livestock. Survival (1 study): One site comparison study in Belgium and the Netherlands reported that the survival of Glanville fritillary caterpillar nests was similar between grasslands with low and high stocking density of sheep. Condition (1 study): One site comparison study in Belgium and the Netherlands found that after 6–10 days of sheep grazing, fewer Glanville fritillary caterpillar nests were damaged in a grassland with lower stocking density than in a grassland with higher stocking density, but there was no difference after two months. BEHAVIOUR (0 STUDIES)Collected Evidencehttps%3A%2F%2Fconservationevidencejournal.com%2Factions%2F3959https%3A%2F%2Fconservationevidencejournal.com%2Factions%2F3959Sun, 14 Aug 2022 10:37:16 +0100Collected Evidence: Collected Evidence: Reduce grazing intensity on grassland by seasonal removal of livestock Seven studies evaluated the effects on butterflies and moths of reducing grazing intensity on grassland by seasonal removal of livestock. Five studies were in the UK, one was in France and one was a review across Europe. COMMUNITY RESPONSE (3 STUDIES) Richness/diversity (3 studies): Two of three replicated, controlled studies (including one randomized, paired study and one randomized study) in the UK found that upland pasture where cattle were removed in the summer, and silage fields where cattle were not grazed in September, had a similar species richness of butterflies to pasture grazed throughout the growing season and silage fields grazed in September. The other study found that grasslands where cattle were removed in the summer had a greater species richness of butterflies (and other pollinators) than grasslands grazed throughout the summer. POPULATION RESPONSE (7 STUDIES) Abundance (7 studies): Three controlled studies (including two replicated, randomized studies) in the UK found that grasslands where cattle or cattle and sheep were removed in the summer, or sheep were removed in the winter, had a higher abundance of butterflies (and other pollinators) and caterpillars than grasslands grazed throughout the summer or all year. Three replicated, controlled studies (including one randomized study and one paired study) in the UK and France found that upland pasture where cattle were removed in the summer, silage fields where cattle were not grazed in September, and semi-natural grasslands where sheep were removed during the peak flowering period, had a similar abundance of butterflies, burnet moths and caterpillars to pasture grazed throughout the growing season, silage fields grazed in September, and rotationally grazed grassland. One review of studies in Europe reported that reducing grazing intensity benefitted 41 out of 67 butterfly species of conservation concern, but did not distinguish between the seasonal removal of livestock and reducing stocking density. BEHAVIOUR (0 STUDIES)  Collected Evidencehttps%3A%2F%2Fconservationevidencejournal.com%2Factions%2F3960https%3A%2F%2Fconservationevidencejournal.com%2Factions%2F3960Sun, 14 Aug 2022 10:37:27 +0100Collected Evidence: Collected Evidence: Reduce cutting frequency on grassland Six studies evaluated the effects on butterflies and moths of reducing cutting frequency on grassland. One study was in each of Slovakia, the Czech Republic, Hungary, the UK, Germany and Italy. COMMUNITY RESPONSE (3 STUDIES) Richness/diversity (3 studies): Two of three replicated studies (including one randomized, paired, controlled study and two site comparison studies) in the UK, Germany and Italy found that meadows cut once/year had a higher species richness of butterflies (along with other pollinators) and moths than meadows cut two or more times/year. The other study found that meadows cut one, two or three times/year all had a similar species richness of butterflies. POPULATION RESPONSE (5 STUDIES) Abundance (5 studies): Three of five replicated studies (including two randomized, paired, controlled studies and three site comparison studies) in Slovakia, the Czech Republic, Hungary, the UK and Italy and found that meadows cut once/year had a similar abundance of all butterflies, and of meadow brown adults and caterpillars and scarce large blue adults, to meadows cut two or three times/year. The other two studies found that meadows cut occasionally or once/year had a higher abundance of Scotch argus and pollinators (including butterflies) than intensively mown grasslands and meadows cut twice/year. BEHAVIOUR (0 STUDIES)  Collected Evidencehttps%3A%2F%2Fconservationevidencejournal.com%2Factions%2F3961https%3A%2F%2Fconservationevidencejournal.com%2Factions%2F3961Sun, 14 Aug 2022 10:37:38 +0100Collected Evidence: Collected Evidence: Increase grazing intensity or cutting frequency on grassland Five studies evaluated the effects on butterflies and moths of increasing grazing intensity or cutting frequency on grassland. Two studies were in Germany, and one study was in each of the Czech Republic, the USA and Israel. COMMUNITY RESPONSE (1 STUDY) Richness/diversity (1 study): One replicated, site comparison study in Germany found that at intensively grazed sites species richness of adult butterflies and burnet moths was lower than at sites with low or no grazing, but caterpillar richness was lower in intensively grazed and low grazing sites than in sites with no grazing. POPULATION RESPONSE (5 STUDIES) Abundance (5 studies): Four studies (including one replicated, randomized, paired, controlled study, one replicated, site comparison study, and two site comparison studies) in Germany, the USA and Israel found that grasslands managed with more intensive grazing or with grazing in addition to rotational burning had a lower abundance of all adult butterflies, adult burnet moths, adult regal fritillaries and spring webworm caterpillar nests than ungrazed grasslands, lightly or moderately grazed grasslands or rotationally burned grasslands with occasional light grazing. However, one of these studies found that, while intensively grazed sites had lower butterfly and burnet moth caterpillar abundance than ungrazed sites, there was no difference in caterpillar abundance between ungrazed and lightly grazed sites. One before-and-after study in the Czech Republic reported that after increasing the cutting frequency on traditional meadows (under agri-environment scheme prescriptions) the abundance of Danube clouded yellow decreased. BEHAVIOUR (0 STUDIES)Collected Evidencehttps%3A%2F%2Fconservationevidencejournal.com%2Factions%2F3962https%3A%2F%2Fconservationevidencejournal.com%2Factions%2F3962Sun, 14 Aug 2022 10:37:50 +0100Collected Evidence: Collected Evidence: Change type of livestock grazing Four studies evaluated the effects on butterflies and moths of changing the type of livestock grazing. One study was in each of the UK, Sweden, China and France. COMMUNITY RESPONSE (2 STUDIES) Richness/diversity (2 studies): Two replicated studies (including one paired, site comparison study and one randomized, controlled study) in Sweden and France found that semi-natural grasslands grazed by cattle or horses had a greater species richness of butterflies and burnet moths than grasslands grazed by sheep. POPULATION RESPONSE (2 STUDIES) Abundance (2 studies): One of two replicated, randomized, controlled studies (including one paired study) in China and France found that semi-natural grasslands grazed by cattle had a higher abundance of butterflies and burnet moths than grasslands grazed by sheep. The other study found that meadow steppe grazed by cattle, goats or sheep for 1–5 years had a similar abundance of butterflies and moths. BEHAVIOUR (1 STUDY) Use (1 study): One replicated, site comparison study in the UK found that a similar proportion of fen meadows were occupied by marsh fritillary caterpillars whether they were managed by cattle, horse or sheep grazing. Collected Evidencehttps%3A%2F%2Fconservationevidencejournal.com%2Factions%2F3963https%3A%2F%2Fconservationevidencejournal.com%2Factions%2F3963Sun, 14 Aug 2022 10:38:00 +0100Collected Evidence: Collected Evidence: Use mixed stocking Three studies evaluated the effects on butterflies and moths of grazing with mixed livestock. All three studies were in the UK and compared grazing with sheep and cattle to sheep only. COMMUNITY RESPONSE (1 STUDY) Richness/diversity (1 study): One replicated, randomized, controlled study in the UK found that grassland plots grazed at low intensity with sheep and cattle had fewer moth species than plots grazed at low intensity with sheep only. POPULATION RESPONSE (3 STUDIES) Abundance (3 studies): Two of three replicated studies (including two randomized, controlled studies and one paired, site comparison study) in the UK found that grassland plots grazed at low intensity with sheep and cattle had a similar abundance of moth caterpillars and small invertebrates including caterpillars to plots grazed at low intensity with sheep only. One of these studies found that in plots grazed at high intensity, the abundance of small invertebrates including caterpillars was lower in plots with sheep and cattle than in sheep only plots. The other study found that grassland plots grazed at low intensity with sheep and cattle had a lower abundance of adult moths than plots grazed at low intensity with sheep only. BEHAVIOUR (0 STUDIES)  Collected Evidencehttps%3A%2F%2Fconservationevidencejournal.com%2Factions%2F3964https%3A%2F%2Fconservationevidencejournal.com%2Factions%2F3964Sun, 14 Aug 2022 10:38:09 +0100Collected Evidence: Collected Evidence: Use rotational grazing Six studies evaluated the effects on butterflies and moths of using rotational grazing. Three studies were in France, two were in the USA and one was in the UK. COMMUNITY RESPONSE (5 STUDIES) Community composition (1 study): One replicated, site comparison study in the USA found that rotational, cattle-grazed grasslands had a similar butterfly community to continuously grazed or patch-burned grasslands. Richness/diversity (5 studies): Two of four replicated studies (including one randomized, controlled study, two controlled studies and one site comparison study) in France and the USA found that rotational cattle- and sheep-grazed grasslands had a greater species richness of butterflies and burnet moths than continuously grazed or patch-burned grassland. Another of these studies found that rotationally sheep-grazed grassland had a similar species richness of butterflies and burnet moths to continuously grazed grassland. The other study found that rotational cattle-grazed pastures had a greater species richness of butterflies than constantly grazed pastures but only at a high, not low, stocking density. One replicated, site comparison study in the USA found that rotationally managed grasslands, including some rotationally grazed grasslands, which were last managed longer ago had a higher species richness of butterflies than more recently managed grasslands. POPULATION RESPONSE (6 STUDIES) Abundance (6 studies): Two of four replicated, controlled studies (including two randomized studies and one paired study) in France and the UK found that rotational cattle- and sheep-grazed grassland had a higher abundance of butterflies and burnet moths and caterpillars (along with other invertebrates) than continuously grazed grasslands. However, one of these studies only found this in the first of three years of management. Another of these studies found that rotationally sheep-grazed grassland had a similar abundance of butterflies and burnet moths to continuously grazed grassland. The other study found that rotational cattle-grazed pastures had a higher abundance of butterflies than constantly grazed pastures but only at a high, not low stocking density. One replicated, site comparison study in the USA found that the abundance of two species was higher, two species were lower and the other five species did not differ in rotationally cattle-grazed grasslands compared to continuously grazed or patch-burned grasslands. One replicated, site comparison study in the USA found that rotationally managed grasslands, including some rotationally grazed grasslands, which were last managed longer ago had a higher abundance of butterflies than more recently managed grasslands. BEHAVIOUR (0 STUDIES)Collected Evidencehttps%3A%2F%2Fconservationevidencejournal.com%2Factions%2F3965https%3A%2F%2Fconservationevidencejournal.com%2Factions%2F3965Sun, 14 Aug 2022 10:38:22 +0100Collected Evidence: Collected Evidence: Use rotational mowing Ten studies evaluated the effects on butterflies and moths of using rotational mowing. Two studies were in each of the USA, the Czech Republic and Switzerland, and one was in each of the UK, Germany, Europe and Japan. COMMUNITY RESPONSE (6 STUDIES) Community composition (1 study): One replicated, site comparison study in the Czech Republic found that grasslands managed with “mixed management”, which included mowing parts of a site at different times and leaving some areas uncut, had a similar community composition of butterflies, but a different community composition of moths, to grasslands managed by complete annual mowing. Richness/diversity (6 studies): Three of four replicated studies (including two paired, controlled studies and two site comparison studies) in Germany, Switzerland, Japan and the Czech Republic found that grasslands managed by mowing strips in alternate years, by mowing and burning one half of the meadow each year, or by mowing parts of a site at different times and leaving some areas uncut, had a greater species richness and diversity of butterflies than grasslands cut in full once/year. However, one of these studies also found that grasslands managed by mowing parts of a site at different times and leaving some areas uncut had a lower species richness of moths than grasslands cut in full once/year. The fourth study found that grasslands managed by leaving a rotational area uncut on each mow had a similar species richness of butterflies and burnet moths to grasslands cut in full twice/year. One replicated, site comparison study in the USA found that rotationally managed grasslands, including some rotationally mown grasslands, which were last managed longer ago had a higher species richness of butterflies than more recently managed grasslands. One replicated, site comparison study in Switzerland found that farms managed with more in-field agri-environment scheme (AES) options, including staggered mowing dates, had a similar species richness of butterflies to farms with fewer AES options. POPULATION RESPONSE (7 STUDIES) Abundance (7 studies): Two replicated, paired, controlled studies (including one randomized study) in Germany and Switzerland found that grasslands managed by mowing strips in alternate years, or by leaving a rotational area uncut on each mow, had a higher abundance of butterflies and burnet moths than grasslands cut in full once or twice per year. One of two replicated, site comparison studies in the USA found that rotationally managed grasslands, including some rotationally mown grasslands, which were last managed longer ago had a higher abundance of butterflies than more recently managed grasslands. The other study found that rotationally mown grasslands had a lower abundance of butterflies in the second year after they were last cut than in the first year after mowing. One replicated, site comparison study in the UK reported that two heath fritillary populations survived on rotationally mown grasslands while six populations went extinct on unmanaged grasslands. One review in Europe reported that rotationally mowing grassland benefitted 27 out of 67 butterfly species of conservation concern. One replicated, site comparison study in Switzerland found that farms managed with more in-field agri-environment scheme (AES) options, including staggered mowing dates, had a similar abundance of butterflies to farms with fewer AES options. BEHAVIOUR (1 STUDY) Use (1 study): One replicated, controlled study in the Czech Republic found that 29 out of 32 butterfly species preferred meadows which were half mown in June and August to meadows cut in full twice/year. The other three species were woodland species which only visited meadows temporarily. Collected Evidencehttps%3A%2F%2Fconservationevidencejournal.com%2Factions%2F3966https%3A%2F%2Fconservationevidencejournal.com%2Factions%2F3966Sun, 14 Aug 2022 10:38:31 +0100Collected Evidence: Collected Evidence: Delay cutting or first grazing date on grasslands to create variation in sward height Seven studies evaluated the effects on butterflies and moths of delaying cutting or first grazing dates on grasslands. Two studies were in Germany and one was in each of the UK, Hungary, Switzerland, Austria and Sweden. COMMUNITY RESPONSE (4 STUDIES) Community composition (1 study): One replicated, site comparison study in Austria found that the community composition of butterflies and day-flying moths was different between early-mown and late-mown grasslands. Richness/diversity (3 studies): One of three replicated, controlled studies (including two randomized studies and two paired studies) in the UK, Germany and Switzerland found that, in one of four years, grassland plots cut once/year in July had a higher species richness of butterflies than plots cut once/year in May. One study found that, in one of three years, grassland strips mulched once/year in September had a lower species richness of butterflies than strips mown once/year after 10 June. The third study found that meadows mown 1–2 times/year after 15 July had a similar species richness of butterflies and burnet moths to meadows mown twice/year after 15 June. POPULATION RESPONSE (6 STUDIES) Abundance (6 studies): Three of four replicated, controlled studies (including three randomized studies and three paired studies) in the UK, Germany, Hungary and Switzerland found that grassland cut once/year in July or September had a greater abundance of butterflies, burnet moths and caterpillars than grassland cut once or twice per year in May or June, but in two of the cases only in one of four or two of three years. The fourth study found that meadows mown once/year in September had a similar abundance of scarce large blue butterflies to meadows mown once/year in May, and abundance remained stable in September-mown meadows but decreased over time in May-mown meadows. One site comparison study in Germany found that a meadow mown once/year after the flight season of tufted marbled skipper had a lower density of eggs than a meadow mown before the flight season. One replicated, site comparison study in Sweden found that meadows where grazing commenced after 15 June (together with a lower stocking density) had a higher abundance of clouded Apollo butterflies than meadows where grazing commenced before 15 June (together with a higher stocking density). BEHAVIOUR (1 STUDY) Use (1 study): One replicated, site comparison study in Austria found that short-tailed blue showed a preference for late-mown meadows, but marbled white and meadow brown preferred early-mown meadows. Collected Evidencehttps%3A%2F%2Fconservationevidencejournal.com%2Factions%2F3967https%3A%2F%2Fconservationevidencejournal.com%2Factions%2F3967Sun, 14 Aug 2022 10:38:44 +0100Collected Evidence: Collected Evidence: Raise cutting height on grasslands Two studies evaluated the effects on butterflies and moths of raising cutting height on grasslands. One study was in each of the UK and Switzerland. COMMUNITY RESPONSE (1 STUDY) Richness/diversity (1 study): One replicated, randomized, controlled study in the UK found that intensively managed grassland plots cut to 10 cm in May and July had a similar species richness of butterflies to plots cut to 5 cm. POPULATION RESPONSE (1 STUDY) Abundance (1 study): One replicated, randomized, controlled study in the UK found that intensively managed grassland plots cut to 10 cm in May and July had a similar abundance of butterflies and caterpillars to plots cut to 5 cm. Survival (1 study): One replicated, randomized, paired, controlled study in Switzerland found that the survival of large white caterpillars in grassland plots cut to 9 cm was similar to in plots cut to 6 cm2. Condition (1 study): One replicated, randomized, paired, controlled study in Switzerland found that a similar proportion of wax model caterpillars were damaged when meadows were cut to 9 cm or 6 cm. BEHAVIOUR (0 STUDIES)  Collected Evidencehttps%3A%2F%2Fconservationevidencejournal.com%2Factions%2F3968https%3A%2F%2Fconservationevidencejournal.com%2Factions%2F3968Sun, 14 Aug 2022 10:38:54 +0100Collected Evidence: Collected Evidence: Use motor bar mowers rather than rotary mowers Two studies evaluated the effects on butterflies and moths of using motor bar mowers rather than rotary mowers. Both studies were in Switzerland. COMMUNITY RESPONSE (1 STUDY) Richness/diversity (1 study): One replicated, site comparison study in Switzerland found that farms managed with more in-field agri-environment scheme options, including using bar mowers instead of rotary mowers, had a similar species richness of butterflies to farms with fewer agri-environment scheme options. POPULATION RESPONSE (2 STUDIES) Abundance (1 study): One replicated, site comparison study in Switzerland found that farms managed with more in-field agri-environment scheme options, including using bar mowers instead of rotary mowers, had a similar abundance of butterflies to farms with fewer agri-environment scheme options. Survival (1 study): One replicated, randomized, paired, controlled study in Switzerland1 found that fewer large white caterpillars were killed when meadows were harvested using a hand-pushed bar mower than with a tractor-pulled rotary mower. Condition (1 study): One replicated, randomized, paired, controlled study in Switzerland found that fewer wax model caterpillars were damaged when meadows were harvested using a hand-pushed bar mower than with a tractor-pulled rotary mower. BEHAVIOUR (0 STUDIES)  Collected Evidencehttps%3A%2F%2Fconservationevidencejournal.com%2Factions%2F3969https%3A%2F%2Fconservationevidencejournal.com%2Factions%2F3969Sun, 14 Aug 2022 10:39:04 +0100Collected Evidence: Collected Evidence: Mark the location of webs or caterpillars before mowing One study evaluated the effects on butterflies and moths of marking the location of webs or caterpillars before mowing. This study was in Poland. COMMUNITY RESPONSE (0 STUDIES)   POPULATION RESPONSE (1 STUDY) Abundance (1 study): One replicated, before-and-after study in Poland reported that after marsh fritillary caterpillar webs were marked before mowing, the number of webs increased the following year. BEHAVIOUR (0 STUDIES)  Collected Evidencehttps%3A%2F%2Fconservationevidencejournal.com%2Factions%2F3970https%3A%2F%2Fconservationevidencejournal.com%2Factions%2F3970Sun, 14 Aug 2022 10:39:27 +0100Collected Evidence: Collected Evidence: Maintain or restore traditional water meadows and bogs Five studies evaluated the effect on butterflies and moths of maintaining or restoring traditional water meadows and bogs. Two studies were in the UK and one was in each of Germany, Belgium and Poland. COMMUNITY RESPONSE (1 STUDY) Richness/diversity (1 study): One replicated, site comparison study in Germany found that the species richness of butterflies was similar in wet meadows managed by mowing once/year in autumn, or by light grazing with cattle or horses. POPULATION RESPONSE (4 STUDIES) Abundance (4 studies): One of two replicated, before-and-after studies in Belgium and Poland found that after cattle grazing on wet grassland began, the number of bog fritillaries which emerged on grazed areas was lower than before grazing started. The other study reported that after trees and shrubs were removed from wet meadows, the number of marsh fritillary caterpillar webs increased the following year. One replicated, site comparison study in Germany found that the abundance of butterflies was similar in wet meadows managed by mowing once/year in autumn, or by light grazing with cattle or horses. One site comparison study in the UK found that wet grassland grazed at an intermediate intensity by cattle had a higher abundance of marsh fritillary caterpillar webs, but a similar abundance of adults, to grassland grazed at high or low intensity. BEHAVIOUR (2 STUDIES) Use (2 studies): One replicated, before-and-after study in Belgium found that after cattle grazing on wet grassland began, the use of the grazed areas by bog fritillaries was lower than before grazing started. One replicated, site comparison study in the UK found that a similar proportion of fen meadows were occupied by marsh fritillary caterpillars whether they were managed by grazing (with cattle, sheep or horses), burning or were unmanaged. Collected Evidencehttps%3A%2F%2Fconservationevidencejournal.com%2Factions%2F3971https%3A%2F%2Fconservationevidencejournal.com%2Factions%2F3971Sun, 14 Aug 2022 10:39:58 +0100Collected Evidence: Collected Evidence: Maintain or restore native wood pasture and parkland We found no studies that evaluated the effects of maintaining or restoring native wood pasture and parkland on butterflies and moths. ‘We found no studies’ means that we have not yet found any studies that have directly evaluated this action during our systematic journal and report searches. Therefore we have been unable to assess whether or not the action is effective or has any harmful impacts. Please get in touch if you know of such a study for this action.Collected Evidencehttps%3A%2F%2Fconservationevidencejournal.com%2Factions%2F3972https%3A%2F%2Fconservationevidencejournal.com%2Factions%2F3972Sun, 14 Aug 2022 10:40:10 +0100Collected Evidence: Collected Evidence: Maintain upland heath/moorland Three studies evaluated the effects of maintaining upland heath/moorland on butterflies and moths. All three studies were in the UK. COMMUNITY RESPONSE (1 STUDY) Richness/diversity (1 study): One replicated, randomized, controlled study in the UK found that lightly grazed or ungrazed upland acid grassland had a higher species richness of moths than commercially grazed grassland. POPULATION RESPONSE (3 STUDIES) Abundance (3 studies): Three controlled studies (including two replicated, randomized studies) in the UK found that ungrazed, lightly grazed or summer grazed upland grassland had a higher abundance of adult moths, moth caterpillars and all caterpillars than grassland grazed at commercial stocking densities or all year round. BEHAVIOUR (0 STUDIES)  Collected Evidencehttps%3A%2F%2Fconservationevidencejournal.com%2Factions%2F3973https%3A%2F%2Fconservationevidencejournal.com%2Factions%2F3973Sun, 14 Aug 2022 10:40:21 +0100Collected Evidence: Collected Evidence: Reduce field size (or maintain small fields) Five studies evaluated the effects of reducing field size on butterflies and moths. Two studies were in Switzerland, and one was in each of Germany, Sweden and the Czech Republic and Poland. COMMUNITY RESPONSE (5 STUDIES) Richness/diversity (5 studies): Two of four replicated, site comparison studies in Sweden, the Czech Republic and Poland and Switzerland found that arable farms (in more diverse landscapes) and landscapes with smaller fields had a higher species richness of butterflies and burnet moths than areas with larger fields. The other two studies found that mixed farms and landscapes with smaller fields had a similar species richness of butterflies to areas with larger fields. One before-and-after study in Germany found that after reducing field size by increasing the length of field edges on a farm, along with increasing the area of meadows and field margins, the species richness of butterflies and burnet moths increased. POPULATION RESPONSE (4 STUDIES) Abundance (4 studies): Four replicated, site comparison studies in Sweden, the Czech Republic and Poland and Switzerland found that arable and mixed farms and landscapes with smaller fields had a higher abundance of butterflies and burnet moths than areas with larger fields. BEHAVIOUR (0 STUDIES)Collected Evidencehttps%3A%2F%2Fconservationevidencejournal.com%2Factions%2F3974https%3A%2F%2Fconservationevidencejournal.com%2Factions%2F3974Thu, 18 Aug 2022 09:04:31 +0100Collected Evidence: Collected Evidence: Manage hedgerows to benefit wildlife (e.g. no spray, gap-filling and laying) Seventeen studies evaluated the effects of managing hedgerows to benefit wildlife on butterflies and moths. Fourteen studies were in the UK, and one was in each of Belgium, Costa Rica and Italy. COMMUNITY RESPONSE (9 STUDIES) Richness/diversity (9 studies): Three replicated, site comparison studies in the UK and Costa Rica found that hedgerows with trees or a more complex structure had a higher species richness or diversity of butterflies and macro-moths than simpler hedgerows without trees. Three of six replicated studies (including three randomized, paired, controlled studies, one randomized, site comparison, and two site comparison studies) in the UK and Italy found that hedgerows cut to allow incremental growth had a higher diversity of caterpillars and pupae than hedgerows cut to the same size, that hedgerows kept between 1–2 m tall had a higher species richness of butterflies than hedgerows kept below 1 m tall and that fields with hedgerows of a larger volume had higher species richness of butterflies than those with hedgerows of a smaller volume, but only in one of two study years. The other three studies found that hedgerows managed according to agri-environment scheme prescriptions (including less frequent or winter cutting, gap-filling and restricted mowing, in one case in combination with other agri-environment scheme habitat) had a similar species richness of butterflies and moths to conventionally managed hedgerows. POPULATION RESPONSE (17 STUDIES) Abundance (17 studies): Four of six replicated studies (including four randomized, paired, controlled studies, one controlled study, and one paired, site comparison study) in the UK found that hedgerows cut once every 2–3 years, cut in autumn, or cut to allow incremental growth, had a higher abundance of adult butterflies and moths, moth caterpillars and pupae and brown hairstreak eggs than hedgerows cut to the same size every winter. However, one of these studies also found that hedgerows cut to allow incremental growth had a similar abundance of moth caterpillars and pupae to hedgerows cut to the same size. The other two studies found that hedgerows managed by gap-filling and cutting every three years had a similar abundance of moths to conventionally managed hedgerows, and that hedgerows cut in winter, or less frequently in autumn, had more concealed moth caterpillars, but a similar abundance of free-living caterpillars, to hedgerows cut annually in autumn. Three of five replicated, site comparison studies (including one paired study) in the UK and Costa Rica found that hedgerows with trees had a similar total abundance of macro-moths to hedgerows without trees. The other two studies found that hedgerows with trees, or with a more complex structure, had a higher abundance of butterflies and pale shining brown moths than simple hedgerows. Two replicated, site comparison studies in Belgium and Italy found that hedgerows managed with scalloped edges, or maintained at below 1 m tall, had more brown hairstreak eggs and a higher abundance of adult butterflies, than hedgerows with straight edges or allowed to grow over 2 m tall. One of two studies (including one controlled and one replicated, site comparison study) in the UK found that laid or coppiced hedgerows had a higher abundance of butterflies than unmanaged hedgerows. The other study found that managed hedgerows had a lower abundance of caterpillars than remnant hedgerows. One replicated, randomized, site comparison study in the UK found that butterfly abundance was higher in fields with hedgerows of a larger volume, but only in one of two study years. One replicated, site comparison study in the UK found that field margins next to hedgerow trees had a higher abundance of most shrub- and tree-feeding, but not grass- and herb-feeding, moth species than margins away from hedgerow trees. BEHAVIOUR (0 STUDIES)Collected Evidencehttps%3A%2F%2Fconservationevidencejournal.com%2Factions%2F3975https%3A%2F%2Fconservationevidencejournal.com%2Factions%2F3975Thu, 18 Aug 2022 09:18:48 +0100Collected Evidence: Collected Evidence: Plant new hedges Seven studies evaluated the effects of planting new hedges on butterflies and moths. Five studies were in the UK and one was in each of Ireland and Canada. COMMUNITY RESPONSE (5 STUDIES) Richness/diversity (5 studies): Three of four site comparison studies (including three replicated and three paired studies) in the UK, Ireland and Canada found that established hedgerows had a higher species richness of butterflies and macro-moths than in-field beetle banks, crops or pasture. The other study found that hedgerows had a similar species richness of butterflies to grass banks between fields. One replicated study in the UK found that gorse, oak and blackthorn planted within hedgerows had more species of arthropods, including caterpillars, than more commonly planted hawthorn. POPULATION RESPONSE (6 STUDIES) Abundance (6 studies): Five of six studies (including one replicated, controlled study, three paired, site comparison studies and two site comparison studies) in the UK, Ireland and Canada found that the abundance of butterflies, moths, macro-moths and gatekeepers was higher along hedgerows than on beetle banks, grass margins without hedgerows, in field interiors, or 5–10 metres away from hedgerows.The other study found that the abundance of butterflies along hedgerows was similar to grass banks between fields without hedgerows. BEHAVIOUR (1 STUDY) Behaviour change (1 study): One site comparison study in the UK found that moths recorded close to hedgerows were more likely to be flying parallel to it than moths recorded further away. Collected Evidencehttps%3A%2F%2Fconservationevidencejournal.com%2Factions%2F3976https%3A%2F%2Fconservationevidencejournal.com%2Factions%2F3976Thu, 18 Aug 2022 10:00:38 +0100Collected Evidence: Collected Evidence: Manage ditches to benefit butterflies and moths We found no studies that evaluated the effects on butterflies and moths of managing ditches to benefit butterflies and moths. ‘We found no studies’ means that we have not yet found any studies that have directly evaluated this action during our systematic journal and report searches. Therefore we have been unable to assess whether or not the action is effective or has any harmful impacts. Please get in touch if you know of such a study for this action.Collected Evidencehttps%3A%2F%2Fconservationevidencejournal.com%2Factions%2F3977https%3A%2F%2Fconservationevidencejournal.com%2Factions%2F3977Thu, 18 Aug 2022 10:39:00 +0100Collected Evidence: Collected Evidence: Protect in-field trees One study evaluated the effects of protecting in-field trees on butterflies and moths. The study was in Sweden. COMMUNITY RESPONSE (1 STUDY) Richness/diversity (1 study): One replicated, site comparison study in Sweden found that where more trees and trees of more species had been retained in pastures, butterfly species richness was higher, but richness was lower when a high proportion of those trees were large. POPULATION RESPONSE (0 STUDIES) BEHAVIOUR (0 STUDIES)Collected Evidencehttps%3A%2F%2Fconservationevidencejournal.com%2Factions%2F3978https%3A%2F%2Fconservationevidencejournal.com%2Factions%2F3978Thu, 18 Aug 2022 10:40:32 +0100Collected Evidence: Collected Evidence: Plant in-field trees (e.g. copses) We found no studies that evaluated the effects of planting in-field trees on butterflies and moths. ‘We found no studies’ means that we have not yet found any studies that have directly evaluated this action during our systematic journal and report searches. Therefore we have been unable to assess whether or not the action is effective or has any harmful impacts. Please get in touch if you know of such a study for this action.Collected Evidencehttps%3A%2F%2Fconservationevidencejournal.com%2Factions%2F3979https%3A%2F%2Fconservationevidencejournal.com%2Factions%2F3979Thu, 18 Aug 2022 10:41:45 +0100Collected Evidence: Collected Evidence: Provide or retain set‐aside areas in farmland Nine studies evaluated the effects of providing or retaining set-aside areas in farmland on butterflies and moths. Three studies were in the UK, and one was in each of Germany, Ireland, Switzerland, Hungary, Finland and the USA. COMMUNITY RESPONSE (5 STUDIES) Community composition (1 study): One replicated, site comparison study in Germany found that butterfly communities in older set-aside fields included species which were less migratory, spent longer as caterpillars, and had fewer generations/year than species found in newer set-aside fields. Richness/diversity (5 studies): Three of four replicated studies (including one randomized, controlled study and three site comparison studies) in Germany, Ireland, Hungary and Finland found that sown or naturally regenerating set-aside had a greater species richness of butterflies and day-flying moths than cereal fields or pasture, especially when the set-aside was sown with less competitive grasses. One of these studies found a higher species richness of butterflies and day-flying moths in second-year set-aside than in first-year set-aside, but another found no difference in butterfly species richness between 1–3-year-old set-aside. The other study found that set-aside fields had a similar species richness of butterflies and moths to arable and pasture fields. One replicated, site comparison study in the USA found that set-aside fields had a similar species richness of butterflies to native prairies. POPULATION RESPONSE (8 STUDIES) Abundance (8 studies): Two of five replicated studies (including one randomized, controlled study and four site comparison studies) in the UK, Ireland, Hungary and Finland found that the abundance of butterflies and day-flying moths in sown set-aside was higher than in cereal fields, especially when the set-aside was sown with less competitive grasses. One of these studies found a higher abundance of butterflies and day-flying moths in second-year set-aside than in first-year set-aside, but another found no difference in butterfly abundance between 1–3-year-old set-aside. The other three studies found that fallow and stubble set-aside had a similar abundance of adult butterflies and butterfly and moth caterpillars to arable fields and pasture. Two site comparison studies (including one replicated study) in the UK and Switzerland found that set-aside fields had a similar abundance of butterfly and moth adults and caterpillars to uncultivated field boundaries and extensively farmed land. One replicated, site comparison study in the USA found that set-aside fields had a similar abundance of butterflies to native prairies in their first year, but a lower abundance of butterflies thereafter. BEHAVIOUR (0 STUDIES)Collected Evidencehttps%3A%2F%2Fconservationevidencejournal.com%2Factions%2F3980https%3A%2F%2Fconservationevidencejournal.com%2Factions%2F3980Thu, 18 Aug 2022 10:53:43 +0100Collected Evidence: Collected Evidence: Create uncultivated margins around intensive arable or pasture fields Ten studies evaluated the effects on butterflies and moths of creating uncultivated margins around intensive arable or pasture fields. Six studies were in the UK, two were in Sweden, and one was in each of Finland and Germany. COMMUNITY RESPONSE (9 STUDIES) Richness/diversity (9 studies): Two of five studies (including four replicated, one randomized, one paired, two controlled and two site comparison studies) in Sweden, the UK and Finland, found that uncultivated margins had a lower species richness or diversity of butterflies than margins sown with grasses and non-woody broadleaved plants (forbs) or wildflowers. One other study found that the species richness of butterflies and day-flying moths was higher in permanent uncultivated margins than in sown fallow plots, and the other two found that the species richness of butterflies and moths was similar in uncultivated and sown margins. Three replicated studies (including one randomized, controlled study and two site comparison studies) in the UK and Germany found that uncultivated margins which were not grazed or cut, or were only cut in spring or autumn, had a higher species richness of butterflies than margins which were cut in summer. Two site comparison studies (including one replicated study) in the UK and Germany found that the species richness of butterflies was higher in longer or wider uncultivated margins than in shorter, narrower or conventional width margins. One of two replicated studies (including one controlled study and one site comparison study) in the UK and Finland found that uncultivated margins had a higher species richness of butterflies and day-flying moths than cereal fields, but the other found that the species richness of butterflies was similar between regenerating margins and cropped field edges. One replicated, paired, site comparison study in Sweden found that uncultivated margins had a higher species richness of butterflies and burnet moths if they were located closer to existing grassland. POPULATION RESPONSE (9 STUDIES) Abundance (9 studies): Six of seven studies (including six replicated, two randomized, four controlled and three site comparison studies) in Sweden, the UK and Finland found that the abundance of butterflies and moths, and of adult but not caterpillar meadow brown, was lower in uncultivated margins than in margins sown with grasses, or grasses and non-woody broadleaved plants (forbs) or wildflowers, or a mixture of grasses and wildflowers. However, one of these studies found that uncultivated margins had similar abundance of butterflies to margins sown with grasses or cereal crop. The other study found that the abundance of butterflies and day-flying moths was higher in permanent uncultivated margins than in sown fallow plots. Two of three replicated, site comparison studies (including two randomized studies) in the UK found that uncultivated margins which were not cut, or were only cut in spring and autumn, had a higher abundance of butterflies, and adult but not caterpillar meadow brown, than margins cut in summer. The other study found that margins which were not cut and grazed had a similar abundance of butterflies to margins which were cut and grazed. Two replicated studies (including one controlled study and one site comparison study) in the UK and Finland found that uncultivated margins had a higher abundance of butterflies and day-flying moths than cereal fields or cropped field edges. One site comparison study in the UK found that the abundance of butterflies in wide uncultivated margins was higher than in conventional margins. One replicated, paired, site comparison study in Sweden found that uncultivated margins had a higher abundance of butterflies and burnet moths if they were located closer to existing grassland. BEHAVIOUR (0 STUDIES)Collected Evidencehttps%3A%2F%2Fconservationevidencejournal.com%2Factions%2F3981https%3A%2F%2Fconservationevidencejournal.com%2Factions%2F3981Thu, 18 Aug 2022 11:06:18 +0100
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What Works in Conservation

What Works in Conservation provides expert assessments of the effectiveness of actions, based on summarised evidence, in synopses. Subjects covered so far include amphibians, birds, mammals, forests, peatland and control of freshwater invasive species. More are in progress.

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